Xinjiang Key Laboratory of Environmental Pollution and Bioremediation, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi, China. School of Ecological and Environmental Sciences, East China Normal University, Shanghai, China. Geomicrobiology Group, School of Life Sciences, University of Dundee, Dundee, Scotland, UK. Xinjiang Key Laboratory of Environmental Pollution and Bioremediation, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi, China; Geomicrobiology Group, School of Life Sciences, University of Dundee, Dundee, Scotland, UK.